LEADER 00000nam 2200313 4500
001 AAI3184729
005 20061206115449.5
008 061206s2005 eng d
020 9780542263118
035 (UnM)AAI3184729
040 UnM|cUnM
100 1 Ledeboer, Nathan Allen
245 10 Sticking around: The role of fimbriae, cellulose, and
colanic acid in biofilm formation by Salmonella enterica
serovar Typhimurium on epithelial cell surfaces
300 186 p
500 Source: Dissertation Abstracts International, Volume: 66-
08, Section: B, page: 4076
500 Supervisor: Bradley D. Jones
502 Thesis (Ph.D.)--The University of Iowa, 2005
520 Salmonella enterica serovar Typhimurium is a Gram-negative
human pathogen that colonizes and invades the intestinal
mucosa. We and others have demonstrated that S.
Typhimurium attaches to and forms biofilm on eukaryotic
cell surfaces in a type 1 fimbriae-dependent manner. To
investigate the effect of biofilm on gene expression, we
compared gene expression between planktonic and biofilm
growth. Microarray results indicated that genes encoding
products involved in antibiotic resistance, central
metabolism, conjugation, intracellular survival, membrane
transport, and fimbrial biosynthesis are significantly
altered by growth in a biofilm. The identification of five
fimbrial gene clusters was of particular interest because
their roles in adherence and intercellular adherence.
Mutations in the bovine colonization factor ( bcf), long
polar (lpf), plasmid encoded ( pef), thin aggregative
(csg), and Sth ( sth) fimbriae were constructed in S.
Typhimurium and the mutants were found to have various
biofilm phenotypes on plastic, HEp-2 cells and chicken
intestinal tissue. The pef and csg mutants were defective
for biofilm formation on each of the surfaces tested,
while the lpf mutant exhibited a complete loss in ability
to form biofilm on chicken intestinal tissue but only an
intermediate loss of biofilm formation on HEp-2 cells and
plastic. The bcf mutant formed increased biofilm on both
HEp-2 cells and chicken intestinal epithelium while the
sth mutant had no detectable biofilm defects
520 In addition, previous work on Salmonella exopolysaccharide
in biofilm indicated that the EPS composition can vary
based upon the substratum on which the bacterial biofilm
forms. We have investigated the role of genes important in
the production of colanic acid and cellulose, common
components of EPS. A mutation in the colanic acid
biosynthetic gene, wcaM, disrupted biofilm formation on
HEp-2 cells and chicken intestinal tissue, although
biofilm formation on plastic was unaffected. A mutation in
the cellulose biosynthetic gene, yhjN, disrupted biofilm
formation on HEp-2 cells, chicken intestinal epithelium,
and on plastic. Our data indicate that Salmonella
attachment to, and growth on, eukaryotic cells represent
complex interactions that are facilitated by species of
EPS and fimbriae. Understanding these interactions is
likely to provide new insights for intervention strategies
for Salmonella colonization and infection
590 School code: 0096
590 DDC
650 4 Biology, Microbiology
690 0410
710 20 The University of Iowa
773 0 |tDissertation Abstracts International|g66-08B
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